The major biotic regions of the southwestern part of North America, which provide the local context of the desert biomes, are show in the map below; the deserts themselves are shown in greater detail in other maps later on. A more detailed consideration of Arizona's biotic communities will be found here.
In the northeastern corner we see the southwestern edge of the great
plains grasslands as they reach into eastern New Mexico. At top right
are two tongues of the Petran (Rocky Mountain)
forest system where they stretch into northern N.M.; a final outlier
of this montane forest is seen in the Sacramento Mtns. of southern N.M.
at centre-right (although similar forests are found at higher elevations
south into Mexico.) The lower mountains of Mexico support forests
and evergreen woodlands which are structurally rather like the Petran forests,
but are floristically distinct - this is the Madrean
system dominating eastern Sonora and western Chihuahua. Small outliers
of this mainly Mexican system extend northwards into the U.S.A., in s.e.
Arizona: the Chiricahua, Huachuca and Patagonia ranges are prominent
examples. The high Mogollon plateau runs s.e. - n.w. across central
Arizona, and on its top it carries Petran montane forest and woodland,
with Madrean woodland and other evergreen systems at lower elevations -
the Interior Arizonan. At lower altitudes
this is represented by a chaparral bush-scrub system. South of the
Madrean appears the Sinaloan system - a dry-deciduous
thorn-woodland vegetation, closely related to that of the adjacent Sonoran
desert. Forrest Shreve, a pioneer biologist of these regions (see later, below) included part of it within the Sonoran desert.) Finally,
of the non-desert systems, we see the Californian,
which here is represented by coastal chaparral scrub, and the Sierran,
which is a montane forest, very like the Petran system.
Here we see the entire North American desert system. We shall
encounter the Great Basin
cold desert mainly while in transit to and from the warm desert systems;
likewise, we spend little or no time in the Chihuahuan
desert. We usually spend a little time in the southern and eastern
margins of the Mojave
desert. The Sonoran desert
is our primary concern, and you can see that most of it is located in Sonora
and Baja California. Later in these notes you will find a table outlining
the major characteristics of these four desert systems, followed by a similar,
more detailed characterisation of the various sub-sections of the Sonoran
desert itself.
Great Basin Elevation largely >1200m. with mountains up to 3000m. Basin & Range topography w/ alkali flats or dry lakes in largely closed basins. Hard frosts >1 week common. Mean precipitation (2-300mm.) +/- evenly distributed throughout year. Most winter precip. as snow.
Simple vegetation - +/- pure stands of low bushes - Atriplex in more alkaline, finer, soils near centre of basins, Artemisia in coarser marginal soils. Few annuals. Cacti , agaves & yuccas unimportant element of flora - few can withstand long, hard frosts.
Mojave Mainly 600-1200m. Basin & range, but mountains lower & more numerous than in Gt, Basin, few supporting other than desert scrub. Winter and spring rain. More arid than Gt. basin, only the margins receiving on average >120mm; mostly ~100mm. or less.
Predominantly shrubby vegetation - ~70% is Larrea & Ambrosia. Few cacti. Few trees, restricted to washes and higher elevations. Prominent here is the endemic Joshua Tree, Yucca brevifolia. Rich winter annual flora. In many ways, a transition between Sonoran & Gt. Basin assemblages.
Sonoran Mostly <600m; entirely <950m. Plains dotted w/ hills & mountains; few closed basins; some permanent rivers. Rains often light and uncertain, bi-seasonal. Lower elevation mean <200mm.; higher elevations get 2-300mm.
Flats dominated by shrubs, as in Mojave. Bajadas, hills & lower slopes have highly diverse life-forms, including many cacti and trees, forming up to 50% of cover, sometimes forming layered vegetation. Higher slopes show Yucca, Nolina, Agave, Dasylirion.
Chihuahuan Entirely east of continental divide. Over 50% >1200m. Vast plains & high mountain ranges. 5-10°C cooler than Sonoran; frosts common in northern sections. 60-70% of precipitation (mean 2-300mm.) in summer months, mostly as rain, but some snow in winter.
Mostly shrubby vegetation, but some few trees mostly in rocky or riparian areas. Cacti small & insignificant. Most obvious plants are Yucca, Dasylirion, Nolina & Agave (A. lechugilla is a marker.) Fouquieria also widespread. Of the very common and widespread spp., only Larrea, Prosopis & Fouquieria in common w/ Sonoran.
Here is a map showing the geographical distribution of precipitation in the Sonoran Desert, with some climate diagrams showing how its seasonal patterning changes on moving east from the Pacific coast.
Notice the especially dry portions of the desert in the Colorado valley
and in the coastal parts of Baja California and Sonora; the generally
smooth increase in precipitation as one moves away from the coasts;
and, in the climate diagrams at the top of the map (see
explanation
of climate diagrams in the Climate & Deserts document),
note the shift from a winter-rain regime in the west (San Diego) through
bi-modal patterns in central Arizona (Gila Bend and Tucson - though Tucson
has more summer than winter rain) to a largely summer rain system in Hermosillo.
Further east still, say in New Mexico and Chihuahua, there is practically
no winter rain atall. This longitudinal shift in rainfall has important
impacts on the vegetation.
Since the graphic isn't entirely clear, here's the key to rainfall levels:
A = 0-50mm; B = 50-100mm; C = 100-200mm; D = 200-300mm;
E = 300-400mm; F = 400-500mm; G = 500-600mm; H = 600-700mm.
The total annual rainfall map above shows only part of the picture that concerns the vegetation. Also of considerable importance are: regularity & seasonality of precipitation, annual means, seasonal extremes & variability in temperature, as well as features of the local soils, topography and so forth. In this diagram we emphasise the substantial climatic differentiation among the various subsections (see below for descriptions of the subsections) of the desert; it plots temperature and rainfall data from the summer and winter three-month periods. The regional means for the winter and summer periods are connected for ease of comparison of the regions and comprehension of the overall pattern.
Our primary focus is the Sonoran desert, which occupies some 300,000 km2, almost 2/3 of which is in Mexico. It extends from 35°N, just north of Needles, on the Colorado River, south to 27°N in mainland Mexico, and to 24°N in Baja California Sur. To the east, the Sonoran desert merges into semi-desert grassland and Chihuahuan desert scrub; to the northeast it merges into chaparral scrub; to the northwest it merges into the Mojave desert scrub; to the west, it merges into the chaparral and conifer woodlands of the Californian mountains.
These boundaries towards the east, north and west correspond quite well with the extent of territory within which there is no more than 24 hours of sub-freezing temperatures (within the desert, though there are some localities entirely without frost, other areas have up to four months with a chance of at least some nightly frost.)
To the south, both in mainland Mexico & in Baja California, the vegetation gradually increases in density & stature until it turns into drought-deciduous thornscrub, where no frosts occur, and it is more mesic. The maps and diagrams have adequately shown the characteristic bimodal rainfall pattern, but they fail to convey the very different character of the two sorts of rains. Winter rain is typically rather gentle, widespread, and of several days duration; summer rain is local, short-lived and often violently torrential. Winter rains achieve good penetration; summer rains mostly finish up in washes, as flash-floods.
Virtually the entire flora of this desert is of subtropical origin, and it has developed in this region surprisingly recently. As the glaciers were receding from much of Canada ~8-9000 years ago, conifer woodlands, similar to those still found in n. Arizona, Nevada and Utah, was receding from the area now occupied by the northern Sonoran Desert. The flora is still evidently in flux, with much evolution presumably still going on; the fauna similarly is recently-arrived, and much of it is still broadly distributed across all of the southern arid belt of N. America.
On the next sections is a synopsis of characteristic features of the several subsections, as originally described by Shreve. Following that are some charts showing, in a little more detail, the ecological extents of some of the major elements of the flora. Later still, you will find a chart showing the diversity of morphological body-forms evolved by desert plants, as they were distinguished by Shreve in his studies of the Sonoran desert. Many of these same body plans, or life-forms, are to be found in desert vegetations elsewhere in the world, even though they have been developed convergently from other phylogenetic groups. As an example: true cacti are restricted to the Americas, but cactus-like stem-succulents have been evolved within the Asclepiadaceae & Euphorbiaceae.
The vegetation of the Sonoran desert is extremely diverse, and this is a reflection of its topographic and climatic diversity (see above). This vegetational diversity was codified over 50 years ago by a pioneer in the study of the North American deserts, Forrest Shreve. He divided the desert into seven regions, which we continue to use here. However, other more recent workers, notably Brown and Lowe have made modifications. Nevertheless, as you can see in the maps below, the essence of Shreve's system is retained. below the maps you will find a table wherein the vegetation types and floras of the subsections are characterised.

| PHYSICAL CHARACTERISTICS | VEGETATION CHARACTERISTICS |
| Lower Colorado-Gila
section
Elevation sea level to 400m.
|
Microphyllous Larrea-Ambrosia desert.
Vegetation typically very simple & open, heavily dominated by small to medium-sized bushes. Flats dominated by Larrea tridentata Ambrosia dumosa; Atriplex canescens found in alkaline areas. Typical of washes: Baccharis sarothroides Chilopsis linearis Ambrosia ambrosoides Justicia californica. Dalea spinosa In washes and on slopes: Prosopis juliflora P. pubescens Cercidium floridum Cercidium microphyllum Olneya tesota Condalia lycioides Lycium andersonii Encelia frutescens Castela emoryi Upper bajadas: Ambrosia deltoidea Fouquieria splendens Krameria grayi Echinocereus engelmannii Opuntia echinocarpa |
| Arizona Upland section
Elevations 150-950m. Precipitation Most mesic
section of the true desert, with overall mean ~300mm., strongly biseasonal
(so only short dry seasons), ~50% in summer months.
|
Crassicaulescent Cercidium-Opuntia desert.
Cover mostly 20-60%. Highly diverse: open scrub to quite substantial 'woodland.' In the more dense stands at higher elevations, direct sight-line can drop to <35m. Succulents and green-stemmed plants abundant; both summer & winter ephemerals. Diverse rooting systems, including taproot (Ephedra), taproot + laterals (Larrea, Prosopis, Ambrosia), and subsurface (most cacti.) Flats w/ fine soils dominated by Larrea tridentata. Typical associates: Prosopis juliflora Acacis constricta A. greggii Opuntia arbuscula O. versicolor O. spinosior O.fulgida Typical of washes: Baccharis sarothroides Ambrosia ambrosoides Justicia californica Chilopsis linearis. Also some large trees: Populus fremontii Fraxinus velutina Juglans major Platanus wrightii In washes and on slopes: Prosopis juliflora Cercidium floridum Cercidium microphyllum Olneya tesota Condalia globosa Lycium fremontii Celtis tala Zyziphus obtusifolia Opuntia phaeacantha Cereus gigantea. Upper bajadas: Ambrosia deltoidea Fouquieria splendens Celtis tala Krameria grayi Echinocereus engelmannii Opuntia echinocarpa O. fulgida O. versicolor O. phaeacantha O. leptocaulis Cereus gigantea Acacia constricta A. greggii Simmondsia chinensis Ephedra trifurca Ferocactus wislizenii Encelia farinosa Calliandra eriophylla Krameria grayi Psilostrophe cooperi Jatropha cardiophylla Canotia holacantha |
| Sonoran Plains section
Elevation ~750m. in east; ~100m. in west.
|
Arbosuffrutescent Olneya-Encelia
desert.
Dominated by open stands of low trees, with shrubs and large cacti. Larrea reduced in abundance, especially in south, as trees increase in abundance & stature. Vines become important. Dominant trees: Prosopis juliflora Cercidium floridum Cercidium microphyllum Olneya tesota. Increasingly frequent in the south are: Ipomaea arborescens Acacia willardiana A. cymbispina Cercidium sonorae C. praecox. Other trees include: Bursera microphylla. Bushes and shrubs include: Celtis pallida Condalia lycioides Jatropha cardiophylla Acacia constricta Caesalpinia pumila Lycium brevipes Krameria parvifolia. In southern reaches, Larrea becomes replaced by Forchammeria, Encelia declines, Fouquieria splendens gives way to F. macdougalli, Cereus giqantea gives way to Stenocereus thurberi and Lophocereus schottii. |
| Sonoran Foothills
section
(now = Sinaloan Thornscrub) Elevation from ~1000m. in east to sea level in west. Precipitation
350-500mm., ~75% in summer months.
|
Arborescent Acacia-Prosopis
desert.
Larrea, Simmondsia virtually absent. Mostly a drought-deciduous low woodland of thorny trees and bushes, inceasing in stature & density towards the south (cover - from 50% in n. to 100% in s.) Cacti of much reduced importance, though Stenocereus and Pachycereus obvious, and Rathbunia alamosensis, Opunia fulgida and Lophocereus are common. Prominent bushes & trees: Prosopis juliflora Acacia cymbispina A. farnesiana A. pennatula Bursera odorata Caesalpinia pumila C. sonorae Ceiba acuminata Encelia farinosa Dodonea viscosa Fouquieria mcdougalli F. diguetii Guaiacum coulteri Eysenhardtia orthocarpa Jatropha cardiophylla Ipomaea arborescens Lantana velutina Lysiloma divaricata Mimosa laxia Olneya tesota Sapium biloculare |
| Central Gulf
Coast section
Elevation sea-level to ~150m.
|
Sarcocaulescent Bursera-Jatropha desert.
Mainly widely-spaced shrubs (Larrea, Jatropha, Euphorbia, Fouquieria) and small trees (Cercidium, Olneya, Bursera) separated by bare ground (cover ~10-20%). The more striking elements of the region are the swollen-trunk plants, often with latex or aromatic resins, and the large columnar cacti. Virtually no low shrub element, in contrast to Arizona Upland section (Ambrosia) or to Vizcaino section (Ambrosia, Viguiera.) In Baja, from north to south, there is a shift from dominance by plants typical of the Colorado desert - Larrea tridentata Fouquieria splendens Cercidium microphyllum - to the following: Bursera laxifolia B. hindsiana Jatropha cinerea Fouquieria diguetii. Cacti are represented by a large number of cholla species (Opuntia bigelovii, O. clavellina, O. cholla, O. ramosissima, and O. tesajo) by cardon (Pachycereus pringlei), and by Ferocactus. Boojum Tree (Fouquieria [Idria] columnaris) also present in some localities. Upland sections, in addition to the taxa mentioned may have Pedilanthus macrocarpus Krameria parvifolia Agave sobria Lycium brevipes. Steamways are markedly different, and may support Lysiloma candida Prosopis juliflora Cercidium floridum.
|
| Vizcaíno
section
Elevation sea-level to ~1700m. with ~75% <500m.
|
Sarcophyllous Agave-Ambrosia desert.
Highly diverse, depending on terrain, from uniform low, sparse, scrub to rich, complex mixes of diverse life-forms. Abundant most places are fleshy-leaved plants e.g. Mesembryanthemum, Agave, Dudleya; often prominent are Idria, Pachycereus, Yucca valida, Pachycormus discolor. Several species of Agave and Ambrosia. Perennial grasses +/- absent. In north, ecotone with California chaparral, with Myrtillocactus cochal Euphorbia misera Bergerocactus emoryi. Characteristic northern vegetation: Ambrosia camphorata A. chenopodifolia Agave shawii Viguieria deltoidea Encelia farinosa Fouquieria splendens Simmondsia chinensis Eriogonum fasciculatum Ephedra californica and chollas (Opuntia.) Vegetation more xeric inland, with A. dumosa and Larrea. Gradually, southern taxa dominate, including Idria columnaris Pachycereus pringlei Lophocereus schotii Agave shawii A. cerulata A. deserti Ambrosia magdalenae Viguiera deltoidea Opuntia molesta O. clavellina Yucca valida Pachycormus discolor Machaerocereus gummosus Jatropha cinerea Pedilanthus macrocarpa Fouquieria diguetii Near coast, fog supports dense epiphyte cover of lichens and bromeliads (Tillandsia recurvata) |
| Magdalena
section
(now part of Vizcaino) Elevation sea-level to ~1500m., mostly below 500m.
|
Arbo-crassicaulescent Lysiloma-Machaerocereus
desert.
Shreve erected this section to acknowledge the absence of the bizarre Idria, Pachycormus, Yucca valida and Agave shawii of the Vizcaino, the prominence of large trees, and of several large cacti. Despite the distinct aspect, the two regions are closely related. In the north, the vegetation is much like of the Gulf Coast, including: Larrea Jatropha cuneata J. cinerea Bursera microphylla. The large trees include: Lysiloma candida Prosopis torreyana Prosopis palmeri Cercidium peninsulare Bursera laxiflora Fouquieria diguetii Notable larger cacti include: Machaerocereus gummosus Stenocereus thurberi Pachycereus pringlei Opuntia cholla Lophocereus schotti . On the sandy plains is found the strange Machaerocereus eruca, an endemic mobile creeping cactus. This plains vegetation also includes: Opuntia cholla Encelia farinosa Ambrosia magdalenae Pachycereus pringlei Jatropha cinerea Fouquieria diguetii Larrea Euphorbia magdalenae E. californica. |
Desert Plant Life-forms -
- the diverse adaptive morphological
responses to desert climates
The deserts of the world support some of the most distinctive plant body forms known and the Sonoran Desert provides most of them to the student. One of the foremost early students of the Sonoran Desert was Forrest Shreve, and he provided a categorization-system to bring some order to the diversity of plant morphologies - what he called Life-Forms. The tree-type diagram below illustrates this system.
It begins by recognising the major distinction between Ephemeral and Perennial plants - those that complete an entire life-cyle, followed by death of the recognisable plant body vs. those wherein the plant body persists several years, perhaps to produce seed several times.
Following that dichotomy, the life-forms are broken down further according to which parts of the plant provide the perennating structures — those that remain alive from one growing season to the next, and other aspects of plant habit.
We shall be meeting examples of all these life-forms during the course, and it is one of your tasks to enter a species name into the boxes in this graphic as an example taxon.
With a few exceptions, a given plant taxon is referred to in texts and articles by use of but a single scientific name (genus and species). By contrast, many taxa have several vernacular names. In our case, this is complicated by there being both English ad Spanish common names. Further, the same vernacular name may refer to several distinct taxa!
This adds up to confusion, and you are encouraged to use taxonomic (Latin/Greek) in preference to vernacular names (after all, some taxa don't even have vernacular names.
However, many sources make reference to plants only by vernacular
names and so to reduce your confusion, here's a correspondence for the
major taxa ch you may encounter.
Acacia sp. Acacias espino, huisache, una
de gato
Agave spp. Century plants lechugilla,
amole, maguey, mescal
Allenrolfea Pickleweed, Iodine bush
Ambrosia spp. Bursage, Burrobush, Western ragweed
(Franseria)
Anisacanthus Hummingbird bush chuparosa
Arctostaphylos sp. Manzanita Manzanita
Argemone Pricklepoppy chicalote
Artemisia spp. Sagebrush
Asclepias spp. Milkweed ajamente
Astragalus Locoweed
Atriplex spp. Saltbush,shadscale chamiso
Baccharis
sarothroides Desert broom
glutinosa seepwillow batamote, hierba del
pasmo
Baileya multiradiata Desert marigold
Beloperone californica see Justicia
Berberis spp. Barberry Agritos, algerita
Bursera spp. Elephant tree copal, torote
Caesalpinia Bird of Paradise
Calliandra Fairyduster, Feather Huajillo, mesquitilla,
duster, False mesquite tabardillo, zapotillo,
cabeza de angel
Canotia holacantha Crucifixion Thorn
Carnegiea gigantea Saguaro
Cassia Senna
Castela emoryi Crucifiction Thorn
Ceanothus Deerbush, buckbrush
Ceiba Silk Cotton Tree, pochote
Kapok Tree
Celtis Hackberry Palo blanco, tala
Cercidium
microphyllum Foothills Paloverde palo verde
floridium Blue Paloverde
sonorae palo estribo
praecox palo brea
peninsulare palo de pua
Cercis Redbud, Judastree
Cereus
greggi Night-blooming cereus Reina de la noche
Lemairocereus thurberi organ-pipe cactus pitahaya
duice
Stenocereus
Lophocereus schotti senita senita
Pachycereus cardon cardon
Chilopsis linearis Desert Willow,
Desert Catalpa
Chrysothamnus Rabbitbrush chamisa
Cleome spp. Burrofat
Cocos Coconut palm cocotera
Coleogyne Blackbrush
Condalia Graythorn Crucillo
Cordia parviflora palo de
asta
Cowania Cliffrose, buckbrush,
quinine bush
Dalea
spinosa Smoketree
emoryi Dye bush
Dasylirion Sotol
Dodonaea Hopbush
Dudleya spp. Liveforever Siempreviva
Echinocereus Hedgehog cactus
Encelia farinosa Brittlebush Incienso
Ephedra spp. Mormon Tea canutillo
Eriogonum spp. Buckwheat
Erythea (Brahea)
armata Palma ceniza
brandegeei Fan Palm palma de taco
Erythrina Coral bean, Coral tree chilicote
Eschscholtzia Mexican Goldenpoppy Copa de oro
Euphorbia spp. spurge candelilla
Eysenhardtia kidneywood
Ferocactus sp. Barrel cactus biznaga
Flourensia Tarbush
Forchammeria jito
Fouquieria
splendens ocotillo
macdougalli Tree Ocotillo palo Adan
diguettii Tree Ocotillo palo Adan
Franseria see Ambrcsia
Fraxinus Ash fresno
Gutierrezia Snakeweed
Haplopappus Burroweed, jimmyweed,
Turpentine bush
Hilaria spp. Tobosa grass
Holacantha emoryi Crucifixion thorn amargoso
(Castela emoryi)
Hymenoclea Burrobrush, Cheeseweed
Idria columnaris BooJum Tree Cirio
Ipomaea arborescens Morning Glory Palo
santo
Jacobinia californicae Chuparosa
Jatropha
cardiophylla (cordata) Limber bush Sangre
de Cristo
cinerea lomboy, zapo,
cuneata torote prieto, sangre
de drago
Juglans walnut
Justicia californica Beloperone chuparosa
Koeberlinia spinosa Crucifixion thorn junco
Krameria Ratany
Lantana lantana
Larrea tridentata Creosote bush gobernadora,
hediondilla
Lycium spp. squawberry Tomatillo
Lysiloma palo blanco, manta
Machaerocereus
gummosus pitahaya agria
eruca Caterpillar cactus
Mammillaria spp. Pincushion cacti, Fishhook cacti
Mesembryanthemum Ice plant Flor del sol
Mimosa spp. Sensitive plant, catclaws
Muhlenbergia spp. muhly grass
Nicotiana Tree tobacco
Nolina Beargrass Sacahuista
Olneya tesota Ironwood Palo fierro
Opuntia
(Cylindropuntia) Chollas
Opuntia
(Platyoeuntia) Prickly Pears
Pachycormus discolor Elephant tree copalquin,
torote blanco
Parkinsonia Mexican paloverde, Horse Bean
Pedilanthus macrocarpus candelilla
Phoenix dactylifera Date palm datil
Phoradendron Desert Mistletoe
Platanus Sycamore
Populus Cottonwood, poplar, aspen alamo
Porophyllum gracile Stinkweed yerba del venado
Prosopis spp. Mesquite, screwbean algarrobo,
tornillo
Psilostrophe Paperflower
Rathbunia octopus cactus
Salsola kali Russian thistle, tumbleweed
Salvia Sage, Desert sage
Sambucus Elderberry
Sapium biloculare Mexican jumping bean
Sarcobatus Greasewood
Simmondsia chinensis Jojoba
Sphaeralcea Globemallow
Suadea Seepweed, inkweed, quelite
iodine bush
Tamarix Saltcedar
Tillandsia recurvata Ballmoss heno pequeno
Tiquilia Borage
Vizcainoa geniculata guayacan
Washingtonia Fan palm palma
Yucca
brevifolia Joshua tree
valida Datilillo
elata Soapweed Datil, palmilla
Zizyphus graythorn crucillo